1986, M. O. Woodburne, C. R. Campbell, T. H. V. Rich, N. S. Pledge, 5: Geology, Stratigraphy, Paleoecology, Michael O. Woodburne, W. A Clemens (Editors), Revision of the Ektopodontidae (Mammalia, Marsupialia, Phalangeroidea) of the Australian Neogene, University of California Geological Publications, Volume 131, page 75,
This apparently reflects a general change from a more lacustrine environment in the lower member to a more fluviatile setting in the upper.